• ACS, Dallas (2014)
  • SBS, Atlanta (2015)
  • 2017
  • SBS, Athens
  • OSM, New Orleans
  • Aquatic Sciences, Honolulu
  • AGU, New Orleans
  • 2018
  • SBS, Tallahassee (organizer)
  • OCB Meeting, Woods Hole

  • Goldschmidt, Boston (presentation)
  • LTER Meeting, California
  • 2019
  • Regional Class Research Vessel Workshop, Gulfport
  • GoMOSES, New Orleans
  • 2020
  • OSM, San Diego (presentation)
  • CCELTER Forum (presentation
  • [link])

Journals Reviewed For

  • Ocean Dynamics
  • JGR-Oceans
  • Deep Sea Research II


Peer Reviewed


Wang, S., Kranz, S.A., Kelly, T. B., Song, H., Cassar, N. & Stukel, M.R., Lagrangian studies of net community production: assessing the effect of diel and multi-day non-steady state factors and vertical fluxes (2020), JGRO.[link; pdf]

Kranz, S.A., Wang, S., Kelly, T. B., Stukel, M.R., Goericke, R., Landry, M.R. & Cassar, N., Lagrangian studies of marine production: a multi-method assessment of productivity relationships in the California Current Ecosystem upwelling region (2020), JGRO.[link; pdf]


Kahru, M., Kelly, T. B. & Stukel, M.R., Satellite estimation of the sinking export flux of carbon in the California Current (2019), DSR2. [link; pdf]

Fender, C., Kelly, T. B. & Stukel, M.R., Ecosystem Function in Export Production: an Optimized UVP Algorithm for the California Current (2019), Frontiers in Marine Science. [link; pdf]

Kelly, T. B., Davison, P. C., Goericke, R., Landry M. R., Ohman, M.D. & Stukel, M.R., The Importance Of Diel Vertical Migrations Of Mesozooplankton For Supporting A Mesopelagic Ecosystem: An Inverse Modeling Approach In The California Current (2019) Frontiers in Marine Science. [link; pdf]

Morton, P. L., Landing, W. L., Shiller, A. M., Kelly, T. B., Donat, J. R., De Carlo, E. H. & Shacat, J.,Shelf Inputs and Lateral Transport of Mn, Co, and Ce in the western North Pacific Ocean (2019), Frontiers in Marine Science. [link; pdf]

Stukel, M. R., Ohman, M. D. & Kelly, T. B., Carbon flux attenuation mediated by zooplankton in the shallow twilight zone: Flux feeders, sinking speeds, and the remineralization length scale (in press), Frontiers in Marine Science. [link; pdf]

Stukel M. R., Kelly, T. B., Aluwihare, L. I., Barbeau, K. A., Goericke, R., Krause, J. W., Landry M. R. & Ohman, M. D., The Carbon:234Thorium ratios of sinking particles in the California Current Ecosystem 1: Relationships with plankton ecosystem dynamics (2019), Marine Chemistry. [link; pdf]

Stukel, M. R. & Kelly, T. B.,The Carbon:234Thorium ratios of sinking particles in the California Current Ecosystem 2: A Theoretical Model (2019), Marine Chemistry.


Kelly, T. B., Goericke, R., Kahru, M., Song, H., & Stukel, M.R., CCEII: Spatial and interannual variability in export efficiency and the biological pump in an eastern boundary current upwelling system with substantial lateral advection (2018) DSRI. [link; pdf]

Morrow, R. M., Ohman, M. D., Goericke, R., Kelly, T. B., Stephens, B. M. & Stukel, M. R., CCE V: Primary Production, Mesozooplankton Grazing, and the Biological Pump in the California Current Ecosystem: Variability and Response to El Nino (2018), DSRI. [link; pdf]

Stukel, M. R., Decima, M., Kelly, T.B., A new approach for incorporating 15N isotopic daa into linear inverse ecosystem models with Markov Chain Monte Carlo sampling (2018), PLOS ONE. [link; pdf]


Yanchilina, A., Yelisetti, S., Wolfson-Schwehr, M., Voss, N., Kelly, T.B., Brizzolara, J., Brown, K., Zayac, J., Fung, M., Guerra, M., Coakley, B., Pockalny, R., Exploring methane gas seepage in the California borderlands (2017), EOS. [link]


Tamburini, F., Kelly, T.B., Weerapana, E., Byers, J.A., Paper to Plastics: An Interdisciplinary Research Project in Sustainability (2014), Journal of Chemical Education. [link; pdf]

**By clicking on "pdf" you are requesting a copy of the manuscript from the author. I will promptly provide you with one.

In Prep

Stukel, M. R., Gerard, T., Kelly, T. B., Knapp, A. N., Laiz-Carrion, R., Lamkin, J., Landry, M. R., Malca, E., Selph, K., Shiroza, A., Shropshire, T. A., & Swalethorp, R., Trophic pathways that support larval Bluefin Tuna in the Gulf of Mexico (in prep).

Knapp, A. N., Thomas, R., Stukel, M. R., Kelly, T. B., Landry, M. R., Selph, K. E., Malca, E., Gerard, T., & Lamkin, J., Constraining the sources of nitrogen fueling phytoplankton and food webs in the Gulf of Mexico using nitrogen isotope budgets (in prep).

Kelly, T. B., Kranz, S. A., Song, H., Kahru, M., Ohman, M. D., Goericke, R., & Stukel, M. R., Biogeochemical Changes 2: Vertical and Horizontal Export of Particulate Organic Carbon in a Sub-Mesoscale Filament (in prep).

Kelly, T. B. Goericke, R., Kranz, S.A., Landry, M.R., Song, H. & Stukel, M. R. Biogeochemical Changes 1: Temporal Evolution of New Production, Primary Productivity, and Export Production along a Mesoscale Filament (in prep).

Kelly, T. B., Stukel, M. R. & Kranz, S., A Reusable Pipeline for NCP Modeling from Oxygen Argon Measurements (in prep), Geoscientific Model Development.

Kelly, T. B., Knapp, A., Landry, M.R., Selph, K., Thomas, R., & Stukel, M. R. The balance of new and export production in the oligotrophic Gulf of Mexico (in prep).

Hamdan, L.J., Hayes, C.T., Roberts, B.J., ... Kelly, T.B.,, Report of the Gulf of Mexico Regional Class Research Vessel Science Workshop (in prep).

Ebling, A., Kelly, T.B., Bressac, M., Guieu, C. & Landing, W., The Missing Link: Characterizing the sea surface microlayer from a controlled atmospheric deposition event (in prep), L&O.

Kranz, S. A., Kelly, T. B., Wang, S., Stukel, M. R. & Cassar, N., Dissolved O2/Ar indicate rapid spatial and temporal changes in NCP along and across an upwelling filament in the California current ecosystem (in prep).

Yingling, N., Kelly, T. B., Shropshire, T., Selph, K., Landry, M.R., Knapp, A., Kranz, S.A. & Stukel, M. R. Phytoplankton nutrient uptake, size structure and biomass distribution in the spawning region of Atlantic Bluefin Tuna in the oligotrophic Gulf of Mexico (in prep).


  • Oxygen Production App [Github]
  • Oxygen-Argon Analysis Pipeline [Github; DOI]
  • Thorium-234 Analysis Suite
  • Light Quality Simulation [link]
  • 15N Linear Inverse Ecosystem Model [Github]



Field & Laboratory

  • Deckboard and in situ incubations
  • Nitrate, Ammonium Uptake Incubations
  • Thorium-Uranium Disequilibrium
  • Surface Tethered Sediment Traps
  • N2 Fixation Incubations
  • Membrane & Inlet Inlet Mass Spectroscopy (MIMS/EIMS)
  • Ion-Coupled Plasma Mass Spectroscopy (ICP-MS)


  • Linear Inverse Modeling for Ecosystems
  • Regional Ocean Modeling System (ROMS)
  • Offline Lagrangian Particle Tracking (LTRANS)
  • MITgcm
  • GENOME Model
  • Programming: R, FORTRAN, Julia, Matlab, Java, & web